Thanks for all the great questions to consider.
After listening up close (I can't believe I didn't do this earlier) -- BOTH tweeters seem to be completely silent!
They're wired up single-wire, with the gold-plated links installed. If I remove the links and connect the wires only to the upper...
I have a pair of Totem Sky driven by Musical Fidelity M5si. Right from the get-go the sound has been muffled, particularly with vocals. I hear this with vinyl, CDs, and digital sources. I have tried more break-in, probably at least 60 hours at this point, but it does not seem to be improving...